\u4f5c\u696d\u304c\u716e\u8a70\u307e\u3063\u3066\u304d\u305f\u6642\u3001\u6307\u3092\u30dd\u30ad\u30dd\u30ad\u3068\u9cf4\u3089\u3059\u3053\u3068\u306f\u306a\u3044\u3067\u3057\u3087\u3046\u304b\u3002\u4e2d\u306b\u306f\u9996\u3092\u30dd\u30ad\u30dd\u30ad\u3059\u308b\u4eba\u3082\u3044\u308b\u304b\u3068\u601d\u3044\u307e\u3059\u3002\u7b46\u8005\u306f\u300c\u9cf4\u3089\u3059\u3068\u3001\u6307\u304c\u592a\u304f\u306a\u308b\u3088\u300d\u3068\u8a00\u308f\u308c\u7d9a\u3051\u3001\u592a\u304f\u306a\u3089\u306a\u3044\u3088\u3046\u306b\u3001\u5de6\u624b\u306e\u85ac\u6307\u3060\u3051\u306f\u9cf4\u3089\u3055\u306a\u3044\u3088\u3046\u306b\u3057\u3066\u304d\u307e\u3057\u305f\u3002\u30a4\u30f3\u30bf\u30fc\u30cd\u30c3\u30c8\u306e\u691c\u7d22\u5019\u88dc\u3067\u3082\u300c\u6307\u30dd\u30ad\u30dd\u30ad \u592a\u304f\u306a\u308b\u300d\u3068\u51fa\u3066\u304f\u308b\u73fe\u72b6\u304c\u3042\u308b\u4e2d\u3001\u672c\u5f53\u306b\u5909\u5f62\u3059\u308b\u306e\u304b\u3001\u5c02\u9580\u5bb6\u306b\u805e\u304d\u307e\u3057\u305f\u3002
\n\u3010\u5199\u771f\u3092\u898b\u308b\u3011\u6307\u3092\u30dd\u30ad\u30dd\u30ad\u9cf4\u3089\u3059\u3068\u592a\u304f\u306a\u308b\uff1f \u9996\u306b\u306f\u6ce8\u610f\u3057\u306a\u3051\u308c\u3070\u3044\u3051\u306a\u3044\u3053\u3068\u3082
\n\u306a\u305c\u4eba\u306f\u6307\u3092\u9cf4\u3089\u3057\u3066\u3057\u307e\u3046\u306e\u3067\u3057\u3087\u3046\u304b\u3002\u300c\u30dd\u30ad\u30dd\u30ad\u300d\u3068\u3044\u3046\u97f3\u304c\u5c0f\u6c17\u5473\u826f\u304f\u805e\u3053\u3048\u308b\u304b\u3089\u3001\u624b\u6301\u3061\u7121\u6c99\u6c70\u3067\u7121\u610f\u8b58\u306b\u9cf4\u3089\u3057\u3066\u3057\u307e\u3046\u304b\u3089\u2500\u2500\u306a\u3069\u305d\u308c\u305e\u308c\u306e\u7406\u7531\u304c\u3042\u308b\u306f\u305a\u3067\u3059\u3002
\n\u4e95\u5c3b\u6574\u5f62\u5916\u79d1\uff08\u5175\u5eab\u770c\u795e\u6238\u5e02\uff09\u306e\u9662\u9577\u3067\u533b\u5e2b\u306e\u4e95\u5c3b\u614e\u4e00\u90ce\u3055\u3093\u306b\u8a71\u3092\u805e\u304d\u307e\u3057\u305f\u3002
\n\u307e\u305a\u6307\u9cf4\u3089\u3057\u3067\u3059\u304c\u3001\u4e95\u5c3b\u3055\u3093\u306f\u300c\u30dd\u30ad\u30dd\u30ad\u9cf4\u3089\u3059\u306e\u306f\u3001\u6307\u306e\u4ed8\u3051\u6839\u306e\u300e\u7b2c3\u95a2\u7bc0\u300f\u3060\u3068\u601d\u3044\u307e\u3059\u304c\u3001\u3053\u3053\u306e\u5909\u5f62\u306f\u307b\u3068\u3093\u3069\u8d77\u3053\u308a\u307e\u305b\u3093\u3002\u592a\u304f\u3082\u306a\u308a\u307e\u305b\u3093\u3002\u79c1\u306e\u8a3a\u7642\u7d4c\u9a13\u304b\u3089\u3082\u3001\u6587\u732e\u304b\u3089\u3082\u3001\u5909\u5f62\u304c\u8d77\u3053\u308b\u306e\u306f\u3001\u6307\u306e\u5148\u7aef\u306e\u300e\u7b2c1\u95a2\u7bc0\u300f\u3068\u5148\u7aef\u304b\u30892\u3064\u76ee\u306e\u300e\u7b2c2\u95a2\u7bc0\u300f\u304c\u307b\u3068\u3093\u3069\u3067\u3059\u300d\u3068\u8a71\u3057\u307e\u3059\u3002
\n\u305f\u3057\u304b\u306b\u3001\u9cf4\u3089\u3059\u3053\u3068\u304c\u591a\u3044\u306e\u306f\u4ed8\u3051\u6839\u306e\u90e8\u5206\u3067\u3059\u3002
\n\u300c\u6307\u3092\u9cf4\u3089\u3059\u3053\u3068\u304c\u5909\u5f62\u306b\u5f71\u97ff\u3059\u308b\u306a\u3089\u3001\u591a\u304f\u306e\u4eba\u306e\u7b2c3\u95a2\u7bc0\u306b\u5909\u5f62\u304c\u751f\u3058\u308b\u306f\u305a\u3067\u3059\u304c\u3001\u305d\u3046\u306f\u306a\u3063\u3066\u3044\u307e\u305b\u3093\u3002\u6307\u3092\u9cf4\u3089\u3059\u3053\u3068\u81ea\u4f53\u306f\u60aa\u304f\u306a\u3044\u3068\u601d\u3044\u307e\u3059\u300d\u3068\u4e95\u5c3b\u3055\u3093\u306f\u8a71\u3057\u307e\u3059\u3002
\n\u300c\u79c1\u3082\u9cf4\u3089\u3059\u7656\u304c\u3042\u308a\u307e\u3059\u304c\u5909\u5f62\u306f\u8d77\u304d\u305a\u30011\u56de\u3082\u6163\u3089\u3057\u305f\u3053\u3068\u306e\u306a\u3044\u53f3\u306e\u4eba\u5dee\u3057\u6307\u306e\u7b2c1\u95a2\u7bc0\u306b\u5909\u5f62\u304c\u5c11\u3057\u751f\u3058\u3066\u3044\u307e\u3059\u300d\u3068\u3057\u305f\u4e0a\u3067\u3001\u300c\u5f37\u304f\u9cf4\u3089\u3057\u305f\u308a\u3001\u983b\u7e41\u306b\u9cf4\u3089\u3057\u305f\u308a\u3057\u904e\u304e\u306a\u3044\u9650\u308a\u306f\u5927\u4e08\u592b\u3002\u75db\u307f\u304c\u306a\u3051\u308c\u3070\u3001\u6307\u306e\u30dd\u30ad\u30dd\u30ad\u97f3\u306f\u6c17\u306b\u305b\u305aOK\u3067\u3059\u300d\u3068\u6559\u3048\u3066\u304f\u308c\u307e\u3057\u305f\u3002
\n\u305d\u3082\u305d\u3082\u6307\u306f\u3001\u3069\u3046\u3057\u3066\u9cf4\u308b\u306e\u3067\u3057\u3087\u3046\u304b\u3002\u5b9f\u306f\u3001\u30dd\u30ad\u30dd\u30ad\u97f3\u306e\u539f\u56e0\u306f\u307e\u3060\u5b8c\u5168\u306b\u306f\u5206\u304b\u3063\u3066\u3044\u307e\u305b\u3093\u3002
\n2015\u5e74\u306b\u30aa\u30f3\u30e9\u30a4\u30f3\u79d1\u5b66\u96d1\u8a8c\u300cPLOS ONE\u300d\u306b\u63b2\u8f09\u3055\u308c\u305f\u8ad6\u6587\u306b\u3088\u308b\u3068\u3001\u30dd\u30ad\u30dd\u30ad\u9cf4\u308b\u73fe\u8c61\uff08\u5c02\u9580\u7528\u8a9e\u3067\u300c\u30c8\u30ea\u30dc\u30cc\u30af\u30ea\u30a8\u30fc\u30b7\u30e7\u30f3\u300d\uff09\u306f\u3001\u95a2\u7bc0\u3092\u5f15\u3063\u5f35\u3063\u305f\u6642\u306b\u3001\u95a2\u7bc0\u5185\u306e\u5727\u529b\u304c\u4e0b\u304c\u308b\u3053\u3068\u3067\u95a2\u7bc0\u6db2\u306e\u4e2d\u306b\u300c\u6c17\u6ce1\u304c\u751f\u3058\u308b\u300d\u304b\u3089\u8d77\u304d\u308b\u3068\u8a3c\u660e\u3057\u307e\u3057\u305f\u3002\u3053\u308c\u306f\u3001\u6a5f\u68b0\u5de5\u5b66\u306a\u3069\u3067\u554f\u984c\u306b\u306a\u308b\u3053\u3068\u304c\u591a\u3044\u3001\u6db2\u4f53\u4e2d\u306b\u6ce1\u304c\u751f\u3058\u308b\u73fe\u8c61\u300c\u30ad\u30e3\u30d3\u30c6\u30fc\u30b7\u30e7\u30f3\u300d\u3068\u540c\u3058\u539f\u7406\u3060\u3068\u3044\u3044\u307e\u3059\u3002
\n\u5f93\u6765\u306e\u8003\u3048\u3067\u306f\u3001\u6c17\u6ce1\u304c\u751f\u3058\u305f\u5f8c\u306b\u300c\u6c17\u6ce1\u304c\u5f3e\u3051\u308b\u300d\u30bf\u30a4\u30df\u30f3\u30b0\u3067\u9cf4\u308b\u3068\u3055\u308c\u3066\u3044\u307e\u3057\u305f\u304c\u3001\u3053\u306e\u7814\u7a76\u304c\u660e\u3089\u304b\u306b\u3057\u305f\u306e\u306f\u300c\u6c17\u6ce1\u304c\u751f\u3058\u308b\u300d\u30bf\u30a4\u30df\u30f3\u30b0\u3067\u9cf4\u308b\u3068\u3044\u3046\u3053\u3068\u3067\u3057\u305f\u3002
\n\u610f\u8b58\u3057\u3066\u6307\u3092\u9cf4\u3089\u3057\u3066\u307f\u308b\u3068\u3001\u6307\u3092\u3072\u306d\u3063\u305f\u77ac\u9593\u306b\u97f3\u304c\u9cf4\u3063\u3066\u3044\u307e\u3059\u3002
\n\u4e95\u5c3b\u3055\u3093\u306f\u300c\u3082\u3057\u3001\u6c17\u6ce1\u304c\u5f3e\u3051\u3066\u9cf4\u308b\u97f3\u3060\u3068\u3059\u308b\u3068\u3001\u4e00\u77ac\u30bf\u30a4\u30e0\u30e9\u30b0\u304c\u751f\u3058\u308b\u306f\u305a\u3067\u3059\u3088\u306d\u3002\u3051\u308c\u3069\u77ac\u6642\u306b\u9cf4\u3063\u3066\u3044\u308b\u3053\u3068\u3082\u3042\u308a\u3001\u79c1\u3082\u3053\u306e\u8ad6\u6587\u306e\u6a5f\u5e8f\u306f\u6b63\u3057\u3044\u3068\u601d\u3044\u307e\u3059\u300d\u3068\u8a71\u3057\u307e\u3059\u3002
\n\u3055\u3089\u306b\u3001\u4e00\u5ea6\u9cf4\u3089\u3057\u305f\u95a2\u7bc0\u304c\u3057\u3070\u3089\u304f\u9cf4\u3089\u306a\u304f\u306a\u308b\u7d4c\u9a13\u3092\u3057\u305f\u3053\u3068\u304c\u3042\u308b\u4eba\u3082\u3044\u308b\u304b\u3068\u601d\u3044\u307e\u3059\u3002\u3053\u308c\u306b\u3064\u3044\u3066\u4e95\u5c3b\u3055\u3093\u306f\u300c\u95a2\u7bc0\u5185\u306b\u95a2\u7bc0\u6db2\u306e\u84b8\u6c17\u304c\u5145\u6e80\u3057\u3066\u5727\u529b\u304c\u4e0a\u304c\u308a\u3001\u95a2\u7bc0\u3092\u5f15\u3063\u5f35\u3063\u3066\u3082\u5727\u529b\u304c\u4e0b\u304c\u3089\u306a\u3044\u305f\u3081\u306b\u6c17\u6ce1\u304c\u3067\u304d\u306a\u3044\u304b\u3089\u3001\u3068\u8aac\u660e\u3067\u304d\u308b\u3068\u601d\u3044\u307e\u3059\u300d\u3068\u3057\u307e\u3059\u3002
\n\u3082\u3046\u4e00\u3064\u3001\u9996\u306b\u3064\u3044\u3066\u3002\u300c\u9996\u304c\u9cf4\u308b\u539f\u7406\u3082\u6307\u304c\u9cf4\u308b\u539f\u7406\u3068\u540c\u3058\u53ef\u80fd\u6027\u304c\u9ad8\u3044\u300d\u3068\u4e95\u5c3b\u3055\u3093\u3002\u9996\u306e\u9aa8\u306e\u300c\u981a\u690e\uff08\u3051\u3044\u3064\u3044\uff09\u300d\u306e\u5f8c\u308d\u306b\u3042\u308b\u300c\u690e\u9593\uff08\u3064\u3044\u304b\u3093\uff09\u95a2\u7bc0\u300d\u304c\u5f15\u3063\u3071\u3089\u308c\u3066\u9cf4\u3063\u3066\u3044\u308b\u306e\u3067\u306f\u3001\u3068\u63a8\u6e2c\u3057\u3066\u3044\u307e\u3059\u3002\u307e\u305f\u3001\u6307\u306f\u4e00\u5ea6\u9cf4\u3089\u3059\u3068\u3057\u3070\u3089\u304f\u9cf4\u3089\u306a\u3044\u306e\u306b\u3001\u9996\u306f\u300c\u30dd\u30ad\u30dd\u30ad\u30dd\u30ad\u300d\u3068\u6570\u56de\u9cf4\u308a\u307e\u3059\u304c\u3001\u3053\u308c\u306f\u300c\u690e\u9593\u95a2\u7bc0\u304c\u8907\u6570\u500b\u3042\u308b\u305f\u3081\u3068\u601d\u308f\u308c\u307e\u3059\u300d\u3068\u4e95\u5c3b\u3055\u3093\u306f\u8a71\u3057\u307e\u3059\u3002
\n\u5b9f\u306f\u3001\u4e95\u5c3b\u3055\u3093\u81ea\u8eab\u3082\u305a\u3063\u3068\u9996\u3092\u9cf4\u3089\u3057\u3066\u304d\u305f\u305d\u3046\u3067\u3059\u3002
\n\u305d\u306e\u4e95\u5c3b\u3055\u3093\u306f50\u6b73\u306e\u6642\u306b\u8efd\u3044\u8133\u6897\u585e\u3092\u60a3\u3044\u307e\u3057\u305f\u300250\u4ee3\u4ee5\u4e0b\u306e\u8133\u6897\u585e\u306e\u539f\u56e0\u3068\u3057\u3066\u3001\u981a\u690e\u306e\u4e2d\u3092\u901a\u308b\u690e\u9aa8\uff08\u3064\u3044\u3053\u3064\uff09\u52d5\u8108\u306e\u89e3\u96e2\u3067\u3001\u8840\u6813\u304c\u98db\u3076\u3053\u3068\u304c\u591a\u3044\u3068\u3055\u308c\u3066\u3044\u307e\u3059\u3002\u4e95\u5c3b\u3055\u3093\u306e\u5834\u5408\u3082\u305d\u308c\u304c\u539f\u56e0\u3067\u3057\u305f\u3002
\n\u9996\u3092\u904e\u5ea6\u306b\u3072\u306d\u308b\u3053\u3068\u3067\u8840\u6813\u304c\u98db\u3076\u3068\u3044\u3046\u5831\u544a\u304c\u3044\u304f\u3064\u3082\u3042\u308b\u3068\u3044\u3044\u307e\u3059\u304c\u3001\u305d\u306e\u8981\u56e0\u306e\u5168\u3066\u304c\u9996\u30dd\u30ad\u30dd\u30ad\u3060\u3068\u306f\u65ad\u8a00\u3067\u304d\u306a\u3044\u3068\u3044\u3044\u307e\u3059\u3002\u4e95\u5c3b\u3055\u3093\u306f\u3059\u3050\u306b\u56de\u5fa9\u3057\u3001\u3044\u307e\u3060\u306b\u9996\u3092\u9cf4\u3089\u3057\u3066\u3044\u308b\u3068\u3044\u3044\u307e\u3059\u304c\u3001\u5ff5\u306e\u305f\u3081\u306b\u512a\u3057\u304f\u9cf4\u3089\u3059\u3088\u3046\u306b\u5fc3\u304c\u3051\u3066\u3044\u308b\u305d\u3046\u3067\u3059\u3002
\n\u305f\u3060\u3001\u300c\u9996\u30dd\u30ad\u30dd\u30ad\u306f\u3001\u5834\u5408\u306b\u3088\u3063\u3066\u306f\u52d5\u8108\u3092\u50b7\u3064\u3051\u308b\u53ef\u80fd\u6027\u304c\u3042\u308a\u307e\u3059\u300d\u3068\u4e95\u5c3b\u3055\u3093\u3002
\n\u3055\u3089\u306b\u3001\u300c\u4ed6\u4eba\u306b\u9996\u3092\u3072\u306d\u3063\u3066\u3082\u3089\u3046\u306e\u306f\u3084\u3081\u305f\u65b9\u304c\u826f\u3044\u3067\u3059\u300d\u3068\u3057\u307e\u3059\u3002\u539a\u751f\u52b4\u50cd\u7701\u306f1991\u5e74\u306b\u51fa\u3057\u305f\u901a\u77e5\u306e\u4e2d\u3067\u3001\u981a\u690e\u306b\u6025\u6fc0\u306a\u300c\u56de\u8ee2\u4f38\u5c55\u64cd\u4f5c\u300d\u3092\u52a0\u3048\u308b\u7642\u6cd5\u3092\u7981\u6b62\u3059\u308b\u5fc5\u8981\u304c\u3042\u308b\u3068\u3057\u3066\u3044\u307e\u3059\u3002
\n\u25cf\u533b\u696d\u985e\u4f3c\u884c\u70ba\u306b\u5bfe\u3059\u308b\u53d6\u6271\u3044\u306b\u3064\u3044\u3066\uff08\u539a\u751f\u52b4\u50cd\u7701\u30011991\u5e746\u670828\u65e5\u4ed8\u3001\u533b\u4e8b\u7b2c58\u53f7\uff09\u3068\u308a\u308f\u3051\u981a\u690e\u306b\u5bfe\u3059\u308b\u6025\u6fc0\u306a\u56de\u8ee2\u4f38\u5c55\u64cd\u4f5c\u3092\u52a0\u3048\u308b\u30b9\u30e9\u30b9\u30c8\u6cd5\u306f\u3001\u60a3\u8005\u306e\u8eab\u4f53\u306b\u640d\u50b7\u3092\u52a0\u3048\u308b\u5371\u967a\u304c\u5927\u304d\u3044\u305f\u3081\u3001\u3053\u3046\u3057\u305f\u5371\u967a\u306e\u9ad8\u3044\u884c\u70ba\u306f\u7981\u6b62\u3059\u308b\u5fc5\u8981\u304c\u3042\u308b\u3053\u3068
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